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These are immature red blood cells with a nucleus (as opposed to mature red blood cells with a nuclear remnant - Howell Jolly bodies).

Causes include:

  • leukoerythroblastic anaemia
  • haemolysis


  • Walport MJ et al (1993). Connective tissue diseases: advances in diagnosis and management. BJHM, 50, 121-31.

Last reviewed 01/2018